Epithelial cells SC mapping

Introduction

In this script we are going to map myeloid cell subtypes onto the Visium slides.

Libraries

library(Seurat)
library(ggpubr)
library(cowplot)
library(dplyr)
library(ggplot2)
library(stringr)
library(readr)
library(SPOTlight)

Setting parameters

Loading necessary paths and parameters

set.seed(123)
source(here::here("misc/paths.R"))
source(here::here("utils/bin.R"))

"{epithelium}/{plt_dir}" %>%
  glue::glue() %>%
  here::here() %>%
  dir.create(path = ,
             showWarnings = FALSE,
             recursive = TRUE)

"{epithelium}/{robj_dir}" %>%
  glue::glue() %>%
  here::here() %>%
  dir.create(path = ,
             showWarnings = FALSE,
             recursive = TRUE)

Extract sample id and get Donor ID

# sample_id <- params$sample_id
sample_id <- "esvq52_nluss5"
donor_id <- id_sp_df[id_sp_df$gem_id == sample_id, ] %>% dplyr::pull(donor_id)

Load data

We have 4 different datasets that we are going to analyze separately. The spatial data comes from the script 03-clustering/03-clustering_integration.Rmd while the sc data can be found in Ramon’s scRNAseq analysis: /scratch/devel/rmassoni/tonsil_atlas_private/2-DOWNSTREAM_PROCESSING/results/R_objects/processed_seurat_objects/processed_seurat_objects/tonsil_integrated_with_harmony_scrublet_annotated.rds.

sp_obj <- "{clust}/{robj_dir}/integrated_spatial.rds" %>%
  glue::glue() %>%
  here::here() %>%
  readRDS(file = .)

# Single cell data
sc_obj <- "{decon}/{robj_dir}/tonsil_integrated_with_harmony_scrublet_annotated.rds" %>%
  glue::glue() %>%
  here::here() %>%
  readRDS(file = .)

# level 5 Epithelium T Cells
lvl4_epi_obj <- "{epithelium}/{robj_dir}/epithelial_annotated_level_4.rds" %>%
  glue::glue() %>%
  here::here() %>%
  readRDS(file = .)

Analysis

Metadata

We are going to use SPOTlight to deconvolute the spatial spots and map the cell types.

We will start by adding the level5 definitive metadata labels to the full object.

lvl4_metadata <- lvl4_epi_obj@meta.data %>%
  dplyr::select(barcode, annotation_level_3)

table(lvl4_metadata$annotation_level_3)

## 
##        Basal cells              Crypt   FDCSP epithelium      Outer surface Surface epithelium             VEGFA+ 
##                 55                 88                 33                 14                 58                 29

sc_obj@meta.data <- sc_obj@meta.data %>%
  tibble::rownames_to_column("barcode") %>%
  dplyr::left_join(lvl4_metadata, by = "barcode") %>%
  dplyr::mutate(
    # Annotation for SPOTlight
    annotation_deconv = dplyr::if_else(
      is.na(annotation_level_3), annotation_level1, annotation_level_3)
    # Annotation for SPOTlight with condensed names
    # annotation_cons_deconv = dplyr::case_when(
    #   annotation_deconv %in% c("Central Mem PASK-", "Central Mem PASK+") ~ "Central Mem",
    #   annotation_deconv %in% c("Follicular Th CXCL13+CBLB+_0", "Follicular Th CXCL13+CBLB+_1", "Follicular Th CXCL13+CBLB+_2") ~ "Follicular Th CXCL13+",
    #   annotation_deconv %in% c("Central Mem PASK-", "Central Mem PASK+") ~ "Central Mem PASK",
    #   TRUE ~ annotation_deconv)
    ) %>%
  # Remove CD4 T cells not anotated in Level 5 since they have been removed in the process
  dplyr::filter(! annotation_deconv %in% c("epithelial", "unknown")) %>%
  tibble::column_to_rownames("barcode")

Subset to 500 cells per cell type

#### Get cell IDs to subset by cluster ####
keep_ids <- lapply(
  split(sc_obj@meta.data, sc_obj@meta.data[, "annotation_deconv"]),
  function(subdf) {
  # Determine n_sample, if the size of the group is < cl_n use all the group, if not just use cl_n
  n_sample <- dplyr::if_else(nrow(subdf) < 500,
                             as.numeric(nrow(subdf)),
                             as.numeric(500))
  # Subset a random selection of that group and get the identifiers
  tmp_ds <- subdf[sample(seq_len(nrow(subdf)), n_sample), ] %>%
    tibble::rownames_to_column("barcodeID") %>%
    dplyr::pull(barcodeID)
  return(tmp_ds)
}) %>%
  purrr::flatten_chr() # flatten the list into a vector

#### Subset seurat object ####
sc_obj_sub <- sc_obj[, keep_ids]

Marker calculation

Since the normalization was carried out with the entire dataset we are going to do normalization again within this subset

sc_obj_sub <- Seurat::NormalizeData(
  sc_obj_sub,
  normalization.method = "LogNormalize",
  scale.factor = 10000,
  verbose = FALSE) %>%
  Seurat::FindVariableFeatures(., nfeatures = 3000, verbose = FALSE) %>% 
  Seurat::ScaleData(., verbose = FALSE)

1st we need to determine the marker genes for each high-level cluster

markers_lvl1_out <- "{epithelium}/{robj_dir}/markers_lvl1_epith.RDS" %>% 
  glue::glue() %>% 
  here::here()

if (file.exists(markers_lvl1_out)) {
  
  markers_lvl1 <- readRDS(file = markers_lvl1_out)
  
} else {
  #### Extract marker genes from each cluster ####
  Seurat::Idents(object = sc_obj_sub) <- sc_obj_sub@meta.data[, "annotation_level1"]
  markers_lvl1 <- Seurat::FindAllMarkers(
    object = sc_obj_sub, 
    assay = "RNA",
    slot = "data",
    verbose = TRUE, 
    only.pos = TRUE,
    max.cells.per.ident = 200)
  
  saveRDS(object = markers_lvl1, file = markers_lvl1_out)
}
DT::datatable(markers_lvl1, filter = "top")

Now find the markers for CD4 T cells level 5 only between CD4 T cells

markers_lvl4_out <- "{epithelium}/{robj_dir}/markers_lvl4_epithelial.RDS" %>% 
  glue::glue() %>% 
  here::here()

if (file.exists(markers_lvl4_out)) {
  
  markers_lvl4 <- readRDS(file = markers_lvl4_out)
  
} else {
  # lvl4 myeloid cells lvl4_metadata
  lvl4_subs <- subset(
    sc_obj_sub,
    subset = annotation_deconv %in% unique(lvl4_metadata$annotation_level_3))
  
  #### Extract marker genes from each cluster ####
  Seurat::Idents(object = lvl4_subs) <- lvl4_subs@meta.data[, "annotation_deconv"]
  
  markers_lvl4 <- Seurat::FindAllMarkers(
    object = lvl4_subs, 
    assay = "RNA",
    slot = "data",
    verbose = TRUE, 
    only.pos = TRUE,
    max.cells.per.ident = 200) 
  
  saveRDS(object = markers_lvl4, file = markers_lvl4_out)
}

DT::datatable(markers_lvl4, filter = "top")

Join markers from both runs. Since we still want to keep the epithelial cells markers in the lower level markers we will add them to each one.

# Keep markers for major cell types
# gene_vec <- c("BCL6", "PRDM1", "FDCSP")
markers_lvl1_sub <- markers_lvl1 %>% dplyr::filter(cluster != "epithelial") %>%
    dplyr::filter( ( avg_log2FC > 1 | pct.1 - pct.2 > 0.75) &
                   p_val_adj < 0.01 & pct.1 > 0.3)
                   # | gene %in% gene_vec)

# Keep markers for general epithelial T cells
markers_lvl1_epith <- markers_lvl1 %>%
  dplyr::filter(cluster == "epithelial") %>%
  # Filter to get CD4 specific markers
  # We are keeping those genes with a logFC > 1 or a pct.1 - pct.2 > 0.75 which
  # have a p_val_adj < 0.01 & pct.1 > 0.6 while keeping CD4
  dplyr::filter( ( avg_log2FC > 1 | pct.1 - pct.2 > 0.75) &
                   p_val_adj < 0.01 & pct.1 > 0.6)

Add general Epithelial cell markers to the specific cell types

markers_lvl4_update <- lapply(as.character(unique(markers_lvl4$cluster)), function (i) {
  # Filter level 5 clusters
  tmp <- markers_lvl4 %>%
    # dplyr::filter(avg_log2FC > 0.75 | pct.1 - pct.2 > 0.4) %>%
    # dplyr::filter(avg_log2FC > 0.5 | pct.1 - pct.2 > 0.4) %>%
    dplyr::filter(cluster == i) %>%
    # Remove ribosomal genes
    dplyr::filter(!stringr::str_detect(string = gene,
                                      pattern = "^RPS|^RPL"))
    
  # Add general CD4 markers
  # tmp %>%
  #   dplyr::bind_rows(markers_lvl1_epith) %>%
  #   dplyr::mutate(cluster = i)
}) %>%
  dplyr::bind_rows()

DT::datatable(markers_lvl4_update, filter = "top")

Combine level-1 and level-4 markers

all_markers <- dplyr::bind_rows(markers_lvl1_sub, markers_lvl4_update) %>%
  dplyr::arrange(dplyr::desc(avg_log2FC)) %>%
  dplyr::group_by(cluster, gene) %>%
  dplyr::filter(dplyr::row_number() == 1)

DT::datatable(all_markers, filter = "top")

SPOTlight deconvolution

2nd run deconvolution

Seurat::Idents(object = sc_obj_sub) <- as.character(sc_obj_sub@meta.data$annotation_deconv)

spotlight_ls <- spotlight_deconvolution(
  se_sc = sc_obj_sub,
  counts_spatial = sp_obj@assays$Spatial@counts,
  clust_vr = "annotation_deconv",
  cluster_markers = all_markers,
  cl_n = 100,
  hvg = 0,
  ntop = NULL,
  transf = "uv",
  method = "nsNMF",
  min_cont = 0)

## [1] "Preparing Gene set"
## [1] "Normalizing count matrix"
## [1] "Seeding initial matrices"
## [1] "Training..."
## [1] "Time to train NMF model was 16.48mins"
## [1] "Deconvoluting spots"
## 
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Save data

decon_fn <- "{epithelium}/{robj_dir}/spotlight_ls_epithelium.rds" %>%
  glue::glue() %>% 
  here::here()
# if (! file.exists(decon_fn) ) {
saveRDS(
  object = spotlight_ls,
  file = decon_fn)
# } else {
#   spotlight_ls <- readRDS(file = here::here(glue::glue("{epithelium}/{robj_dir}/spotlight_ls_cd4.rds")))
# }

3rd Check Topic profiles

nmf_mod_ls <- spotlight_ls[[1]]
nmf_mod <- nmf_mod_ls[[1]]

h <- NMF::coef(nmf_mod)
rownames(h) <- paste("Topic", 1:nrow(h), sep = "_")
topic_profile_plts <- SPOTlight::dot_plot_profiles_fun(
  h = h,
  train_cell_clust = nmf_mod_ls[[2]])

topic_profile_plts[[2]] +
  ggplot2::theme(axis.text.x = ggplot2::element_text(angle = 90), 
                 axis.text = ggplot2::element_text(size = 12))

Visualize deconvolution results

decon_mtrx <- spotlight_ls[[2]]
decon_mtrx <- decon_mtrx[, colnames(decon_mtrx) != "res_ss"]

# Set as 0 cell types predicted to be under 1 % of the spot
# decon_mtrx[decon_mtrx < 0.03] <- 0

Change column names

nm_df <- data.frame(
  mod_nm = stringr::str_replace_all(string = unique(all_markers$cluster),
                                    pattern = "[[:punct:]]|\\s|\\+" ,
                                    replacement = "."),
  plt_nm = unique(all_markers$cluster)
  )

# Save nm_df
"{epithelium}/{robj_dir}/epithelium_nm_df.rds" %>%
  glue::glue() %>% 
  here::here() %>%
  saveRDS(
    object = nm_df,
    file = .)


# new_cn <- data.frame(mod_nm = colnames(decon_mtrx)) %>%
#   dplyr::left_join(nm_df, by = "mod_nm") %>%
#   # Central.Mem.PASK. fives some trouble because it only changes between + an -
#   # negative goes first and distinct solves it automatically
#   dplyr::distinct() %>%
#   dplyr::pull(plt_nm)
# 
# colnames(decon_mtrx) <- new_cn

We are going to add the deconvolution to the Seurat object.

rownames(decon_mtrx) <- rownames(sp_obj@meta.data)

sp_obj@meta.data <- sp_obj@meta.data %>%
  tibble::rownames_to_column("barcode") %>%
  dplyr::left_join(data.frame(decon_mtrx) %>%
                     tibble::rownames_to_column("barcode"), by = "barcode") %>%
  tibble::column_to_rownames("barcode")

Look at cells topic profile

basis_spotlight <- data.frame(NMF::basis(spotlight_ls[[1]][[1]]))

train_labs <- spotlight_ls[[1]][[2]]
colnames(basis_spotlight) <- unique(stringr::str_wrap(train_labs, width = 30))

basis_spotlight[basis_spotlight < 0.0000001] <- 0

DT::datatable(basis_spotlight, filter = "top")

Cell type location

Look at the location of each cell type in each slice separately

# Iterate over cell types
ct <- colnames(decon_mtrx)
# Seurat::Idents(sp_obj) <- sp_obj@meta.data$gem_id
sp_sub <- sp_obj[, sp_obj@meta.data$gem_id == sample_id]
sp_sub@images <- sp_sub@images[Seurat::Images(sp_sub) == sample_id]

ct_plt_ls <- lapply(ct, function(i) {
  tmp_plt <- Seurat::SpatialFeaturePlot(object = sp_sub,
                                        features = i,
                                        alpha = c(0, 1)) +
    ggplot2::scale_fill_gradientn(
      colors = heat.colors(10, rev = TRUE)) +
    ggplot2::scale_alpha(range = c(0, 1)) +
    ggplot2::labs(title = stringr::str_wrap(string = i,
                                   width = 25),
         fill = "") +
    ggplot2::theme(
      plot.title = ggplot2::element_text(
        hjust = 0.5,
        size = 20,
        face = "bold"))
  
  if (sum(sp_sub@meta.data[, i]) == 0) {
    tmp_plt <- suppressMessages(tmp_plt + ggplot2::scale_alpha(range = c(0,0)))
  }
  
  return(tmp_plt)
})

(plt_arr <- cowplot::plot_grid(
  plotlist = ct_plt_ls,
  axis = "trbl",
  align = "hv",
  nrow = 5,
  ncol = 5))

"{epithelium}/{plt_dir}/cell_type_location_epithelium_{donor_id}.pdf" %>%
  glue::glue() %>%
  here::here() %>%
  cowplot::save_plot(
    filename = .,
    plot = plt_arr,
    base_height = 25,
    base_width = 25)

Session Info

sessionInfo()

## R version 4.0.1 (2020-06-06)
## Platform: x86_64-conda_cos6-linux-gnu (64-bit)
## Running under: Red Hat Enterprise Linux Server release 6.7 (Santiago)
## 
## Matrix products: default
## BLAS/LAPACK: /scratch/groups/hheyn/software/anaconda3/envs/spatial_r/lib/libopenblasp-r0.3.12.so
## 
## locale:
##  [1] LC_CTYPE=en_US.UTF-8       LC_NUMERIC=C               LC_TIME=es_ES.UTF-8        LC_COLLATE=en_US.UTF-8     LC_MONETARY=es_ES.UTF-8    LC_MESSAGES=en_US.UTF-8    LC_PAPER=es_ES.UTF-8       LC_NAME=C                  LC_ADDRESS=C               LC_TELEPHONE=C             LC_MEASUREMENT=es_ES.UTF-8 LC_IDENTIFICATION=C       
## 
## attached base packages:
## [1] parallel  stats     graphics  grDevices utils     datasets  methods   base     
## 
## other attached packages:
##  [1] tidyr_1.1.3         nnls_1.4            edgeR_3.32.1        limma_3.46.0        Matrix_1.3-4        NMF_0.23.0          Biobase_2.50.0      BiocGenerics_0.36.0 cluster_2.1.0       rngtools_1.5        pkgmaker_0.32.2     registry_0.5-1      tibble_3.1.2        purrr_0.3.4         SPOTlight_0.1.7     readr_1.4.0         stringr_1.4.0       dplyr_1.0.7         cowplot_1.1.1       ggpubr_0.4.0        ggplot2_3.3.5       SeuratObject_4.0.2  Seurat_4.0.3       
## 
## loaded via a namespace (and not attached):
##   [1] readxl_1.3.1          backports_1.2.1       plyr_1.8.6            igraph_1.2.6          lazyeval_0.2.2        splines_4.0.1         crosstalk_1.1.1       listenv_0.8.0         scattermore_0.7       gridBase_0.4-7        digest_0.6.27         foreach_1.5.1         htmltools_0.5.1.1     fansi_0.5.0           magrittr_2.0.1        doParallel_1.0.16     tensor_1.5            ROCR_1.0-11           openxlsx_4.2.3        globals_0.14.0        matrixStats_0.59.0    spatstat.sparse_2.0-0 colorspace_2.0-2      ggrepel_0.9.1         haven_2.4.1           xfun_0.26             crayon_1.4.1          jsonlite_1.7.2        spatstat.data_2.1-0   iterators_1.0.13      survival_3.2-11       zoo_1.8-9             glue_1.4.2            polyclip_1.10-0       gtable_0.3.0          leiden_0.3.8          car_3.0-10            future.apply_1.7.0    abind_1.4-5           scales_1.1.1          DBI_1.1.1             rstatix_0.7.0         miniUI_0.1.1.1        Rcpp_1.0.6            viridisLite_0.4.0     xtable_1.8-4          reticulate_1.20       spatstat.core_2.2-0   foreign_0.8-81        DT_0.18               htmlwidgets_1.5.3     httr_1.4.2            RColorBrewer_1.1-2    ellipsis_0.3.2       
##  [55] ica_1.0-2             farver_2.1.0          pkgconfig_2.0.3       sass_0.4.0            uwot_0.1.10           deldir_0.2-3          locfit_1.5-9.4        utf8_1.2.1            here_1.0.1            labeling_0.4.2        tidyselect_1.1.1      rlang_0.4.11          reshape2_1.4.4        later_1.2.0           munsell_0.5.0         cellranger_1.1.0      tools_4.0.1           cli_2.5.0             generics_0.1.0        broom_0.7.6           ggridges_0.5.3        evaluate_0.14         fastmap_1.1.0         yaml_2.2.1            goftest_1.2-2         knitr_1.35            fitdistrplus_1.1-3    zip_2.2.0             RANN_2.6.1            pbapply_1.4-3         future_1.21.0         nlme_3.1-150          mime_0.11             compiler_4.0.1        rstudioapi_0.13       plotly_4.9.4.1        curl_4.3.1            png_0.1-7             ggsignif_0.6.1        spatstat.utils_2.2-0  bslib_0.2.5.1         stringi_1.6.2         highr_0.9             ps_1.6.0              forcats_0.5.1         lattice_0.20-44       vctrs_0.3.8           pillar_1.6.1          lifecycle_1.0.0       spatstat.geom_2.2-0   lmtest_0.9-38         jquerylib_0.1.4       RcppAnnoy_0.0.18      data.table_1.14.0    
## [109] irlba_2.3.3           httpuv_1.6.1          patchwork_1.1.1       R6_2.5.0              promises_1.2.0.1      KernSmooth_2.23-18    gridExtra_2.3         rio_0.5.26            parallelly_1.26.0     codetools_0.2-18      MASS_7.3-54           assertthat_0.2.1      rprojroot_2.0.2       withr_2.4.2           sctransform_0.3.2     mgcv_1.8-36           hms_1.1.0             grid_4.0.1            rpart_4.1-15          rmarkdown_2.11        carData_3.0-4         Rtsne_0.15            shiny_1.6.0